COLUMBUS, Ohio (AP) — The world's first gorilla born in a zoo, a female named Colo who became the oldest known living gorilla in the U.S., has died at age 60, the Columbus Zoo and Aquarium said Tuesday.

Colo died in her sleep overnight, less than a month after her birthday, the zoo announced. She surpassed the usual life expectancy of captive gorillas by two decades.

"Colo touched the hearts of generations of people who came to see her and those that cared for her over her long lifetime," zoo president Tom Stalf said in a statement. "She was an ambassador for gorillas and inspired people to learn more about the critically endangered species and motivated them to protect gorillas in their native habitat."

Colo had been on exhibit in view of visitors on Monday, when the zoo offered free admission for Martin Luther King Jr. Day. Fond of baseball caps, the gorilla was seen in her enclosure toting around a yellow cap with a gorilla on it, given to her by the zoo team caring for her.

The zoo said the gorilla's body will be cremated and the ashes will be buried at an undisclosed location at the zoo.

Colo was born at the zoo on Dec. 22, 1956. She eventually became a mother of three, grandmother of 16, great-grandmother of 12 and great-great-grandmother of three.

Hundreds of people had gathered at the zoo last month to see Colo for her birthday, when the zoo decorated her enclosure with multicolored construction paper chains and cakes of squash and beet and cornbread with mashed potato parsley frosting.

Zoo officials said a post-mortem examination was planned to determine Colo's cause of death. The median life expectancy for female gorillas in human care is 37.5 years.

Veterinarians recently removed a malignant tumor from Colo, but the zoo had said she was doing well. The necropsy could help show whether the cancer contributed to her death, but the results aren't anticipated for several weeks.

Zoo officials said they also planned to take blood and tissue samples to assist with zoologists' efforts to learn more about western lowland gorillas.

Colo is the second zoo gorilla to die in Ohio this month. Cleveland Metroparks Zoo said one of its two male western lowland gorillas, a 32-year-old named Bebac, died Jan. 6.

Zoo experts say animals in human care are living longer than ever as early diagnosis and improvements in medical care extend their lives. The oldest known living male gorilla, Ozzie, is 55 years old and lives at the Atlanta Zoo, which has a geriatric gorilla specialty.

Packy, an Asian elephant at the Oregon Zoo, is now 54 and the oldest male of his species in North America.

Nikko, a 33-year-old snow monkey at the Minnesota Zoo, is the oldest male snow monkey in North America. Zulie, a 30-year-old Black Howler monkey at the Beardsley Zoo in Bridgeport, Connecticut, is the oldest living female Howler monkey in captivity.

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